Learn / Metabolism

Metabolic pathways

Every FMK pathway as a clickable map. Pick one pathway to hammer it — next step, enzyme, regulator, cofactor, and the facts that get tested (NADPH and who needs it, rate-limiting steps, deficiencies) — or set it to all pathways for a mixed drill. Every answer says which enzyme and which pathway.

Pathway
PDH & TCA cycle · Mitochondrial matrix. Enzymes sit beside the arrows; a ★ marks the rate-limiting step. The map scrolls inside its panel.
Zoom
PyruvatePyruvateAcetyl-CoAAcetyl-CoAOxaloacetateOxaloacetateCitrateCitrateIsocitrateIsocitrateα-Ketoglutarateα-KetoglutarateSuccinyl-CoASuccinyl-CoASuccinateSuccinateFumarateFumarateMalateMalateOxaloacetate (regenerated)Oxaloacetate (regenerated)Pyruvate dehydrogenase complex(PDH)Pyruvate carboxylaseCitrate synthaseCitrate synthaseAconitaseIsocitrate dehydrogenase ★α-Ketoglutarate dehydrogenasecomplexSuccinyl-CoA synthetase (succinatethiokinase)Succinate dehydrogenase (ComplexII)FumaraseMalate dehydrogenase

PDH & TCA cycle

Pyruvate → acetyl-CoA (PDH, five cofactors: 'Tender Loving Care For Nancy'), then acetyl-CoA + oxaloacetate → citrate and round the cycle: 3 NADH + 1 FADH₂ + 1 GTP + 2 CO₂ per acetyl-CoA (~10 ATP). Regulated at citrate synthase, isocitrate DH (rate-limiting) and α-KG DH.

Click any metabolite on the map for its reactions. Source: FMK-02.1 Glycolysis & TCA; FMK-02.4 Mitochondrial metabolism.

Reactions

StepEnzymeCofactorsRegulation
Pyruvate → Acetyl-CoA Pyruvate dehydrogenase complex (PDH)
PDH deficiency — neonatal lactic acidosis, neurologic defects; treat with a ketogenic diet (lysine, leucine) so fuel bypasses pyruvate
Pyruvate dehydrogenase complex (PDHC) deficiency →
TPP (thiamine, B₁), Lipoic acid, CoA (pantothenate, B₅), FAD (riboflavin, B₂), NAD⁺ (niacin, B₃)
+ Pyruvate (inhibits PDH kinase); Ca²⁺ & insulin (activate PDH phosphatase → dephosphorylated, active E1)
Acetyl-CoA; NADH; ATP (all activate PDH kinase → phosphorylated, inactive E1); Arsenite (binds lipoic acid)
Pyruvate → Oxaloacetate Pyruvate carboxylase Biotin (B₇), ATP
+ Acetyl-CoA
Acetyl-CoA → Citrate Citrate synthase
ATP; NADH; Citrate (product)
Oxaloacetate → Citrate Citrate synthase
Citrate → Isocitrate Aconitase
Isocitrate → α-Ketoglutarate Isocitrate dehydrogenase NAD⁺ → NADH, CO₂ released
+ ADP; Ca²⁺
ATP; NADH
α-Ketoglutarate → Succinyl-CoA α-Ketoglutarate dehydrogenase complex TPP (B₁), Lipoic acid, CoA, FAD, NAD⁺ → NADH, CO₂ released
+ Ca²⁺
Succinyl-CoA; NADH; ATP; Arsenite
Succinyl-CoA → Succinate Succinyl-CoA synthetase (succinate thiokinase) GDP → GTP (substrate-level)
Succinate → Fumarate Succinate dehydrogenase (Complex II) FAD → FADH₂
Malonate (competitive)
Fumarate → Malate Fumarase
Malate → Oxaloacetate (regenerated) Malate dehydrogenase NAD⁺ → NADH

Conditions that live on this map

Beriberi (wet and dry)Korsakoff syndromeMaple syrup urine disease (MSUD)Pyruvate dehydrogenase complex (PDHC) deficiencyThiamine (vitamin B₁) deficiencyWernicke encephalopathy

Blue pills sit on one specific arrow; grey ones are whole-pathway problems. Each opens the full condition card, which links back here with the arrow lit.

Facts worth knowing

Yield

  • Per acetyl-CoA, one turn of the TCA cycle yields — 3 NADH + 1 FADH₂ + 1 GTP + 2 CO₂ (~10 ATP)
  • Steps of the TCA cycle that release CO₂ — Isocitrate dehydrogenase and α-ketoglutarate dehydrogenase
  • The only TCA enzyme that makes FADH₂ — Succinate dehydrogenase (Complex II)

Cofactor

  • Enzymes that need thiamine (TPP) — PDH, α-ketoglutarate dehydrogenase, branched-chain α-keto acid dehydrogenase, transketolase
  • Cofactor of pyruvate carboxylase — Biotin (B₇) with ATP; activated by acetyl-CoA
  • Mnemonic 'Tender Loving Care For Nancy' stands for — TPP, Lipoic acid, CoA, FAD, NAD⁺ — the PDH / α-KG DH cofactors

Shuttle

  • Cytosolic NADH from glycolysis enters mitochondria via — Malate-aspartate shuttle (2.5 ATP each) or glycerol-3-phosphate shuttle (1.5 ATP each)

Enzyme deficiency

  • Thiamine deficiency blocks which two TCA-related enzymes? — PDH and α-ketoglutarate dehydrogenase → ↑ pyruvate & lactate, ↓ ATP (Wernicke (confusion, ataxia, ophthalmoplegia), Korsakoff, wet/dry beriberi.)
  • Deficiency of Pyruvate dehydrogenase complex (PDH) causes… — PDH deficiency — neonatal lactic acidosis, neurologic defects; treat with a ketogenic diet (lysine, leucine) so fuel bypasses pyruvate (E1 pyruvate decarboxylase (TPP), E2 dihydrolipoyl transacetylase (lipoic acid, CoA), E3 dihydrolipoyl dehydrogenase (FAD, NAD⁺). Thiamine deficiency blocks it → pyruvate → lactate. Give thiamine BEFORE glucose.)

Rate-limiting step

  • Rate-limiting enzyme of pdh & tca cycle — Isocitrate dehydrogenase (First oxidative decarboxylation; the rate-limiting step of the cycle.)
Something went wrong. Reload 🗙